{"id":24840,"date":"2025-05-24T07:32:03","date_gmt":"2025-05-24T05:32:03","guid":{"rendered":"https:\/\/www.meslibellules.fr\/blog\/?page_id=24840"},"modified":"2025-05-26T12:58:34","modified_gmt":"2025-05-26T10:58:34","slug":"anax-imperator","status":"publish","type":"page","link":"https:\/\/www.meslibellules.fr\/blog\/anax-imperator\/","title":{"rendered":"Anax imperator (Leach in Brewster, 1815)"},"content":{"rendered":"\n<p>L&rsquo;Anax imperator (Aeshnidae) est un des plus spectaculaires Odonates d&rsquo;Europe, par son vol dominant, sa tr\u00e8s grande taille (66 \u00e0 84 mm) et l&rsquo;\u00e9l\u00e9gance de sa livr\u00e9e. Il est tr\u00e8s commun en plaine.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><img decoding=\"async\" width=\"1987\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-male-1-accueil.webp\" alt=\"Anax imperator m\u00e2le, le Fuilet (France-49), 12\/07\/2008\" class=\"wp-image-24842 lazyload\" style=\"--smush-placeholder-width: 1987px; --smush-placeholder-aspect-ratio: 1987\/1504;width:514px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-male-1-accueil.webp 1987w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-male-1-accueil-768x581.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-male-1-accueil-1536x1163.webp 1536w\" data-sizes=\"(max-width: 1987px) 100vw, 1987px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><figcaption class=\"wp-element-caption\">Anax imperator m\u00e2le, le Fuilet (France-49), 12\/07\/2008<\/figcaption><\/figure>\n\n\n\n<h5><span style=\"text-decoration: underline;\">Description &#8211; Identification<\/span><\/h5>\n\n\n\n<p><strong>Les m\u00e2les<\/strong> matures de l&rsquo;Anax empereur (1) ont des yeux verts ou bleu-vert et portent <em>un pentagone sur le front<\/em>. Le thorax est vert sans bandes ant\u00e9hum\u00e9rales ni motifs, mais <em>bleu juste en avant des ailes<\/em>, le premier segment est \u00e9galement vert, le reste de l&rsquo;abdomen est bleu avec une bande noire m\u00e9diane.<br>Ils sont souvent en vol un ou deux m\u00e8tres au-dessus de l&rsquo;eau et on peut donc facilement les confondre :<br>&#8211; au printemps avec <a href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/brachytron-pratense\/brachytron-pratense-male-2\/brachytron-pratense-male-5big.jpg\">Brachytron pratense<\/a>, l&rsquo;Aeschne printanni\u00e8re qui semble aussi ne jamais se poser. Il est plus petit, son thorax est marron devant, avec une bande ant\u00e9hum\u00e9rale jaune et deux lignes brunes sur la face lat\u00e9rale du thorax (pas de bleu avant les ailes), son abdomen est plus sombre. <em>Son vol est erratique<\/em>, il zigzague et cherche les femelles le long des rives en volant bas.<br>&#8211; en \u00e9t\u00e9 avec <a href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/Aeshna\/aeshnaaffinis\/aeshna-affinis-accueil\/aeshna-affinis-accueil-big.jpg\">Aeshna affinis<\/a> dont le thorax est marron dessus et <em>jaune sur les c\u00f4t\u00e9s<\/em>.<br><br><a href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax-parthenope\/anax-parthenope-male-1\/anax-parthenope-male-3big.jpg\">Anax parthenope<\/a> est plus rare, plus petit, avec un thorax sombre, des segments deux et partiellement trois bleus, le reste des segments \u00e9tant plus sombre.<br><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/anax-ephippiger-male-2.jpg\">Anax epipphiger<\/a>, aux yeux bruns, est occasionnel, plus terne, couleur sable avec une selle bleue limit\u00e9e au segment 2.<br><br>En juin et juillet, une tr\u00e8s grande libellule vivement color\u00e9e qui arpente incessamment un segment de rive d&rsquo;une grande mare ou d&rsquo;un \u00e9tang, en chassant les intrus ou les adversaires, a de tr\u00e8s fortes chances d&rsquo;\u00eatre notre Empereur.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-femelle-accueil.webp\"><img decoding=\"async\" width=\"1692\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-femelle-accueil.webp\" alt=\"Anax imperator femelle, La Croix Fourreau (France-49), 25\/08\/2019\" class=\"wp-image-24861 lazyload\" style=\"--smush-placeholder-width: 1692px; --smush-placeholder-aspect-ratio: 1692\/1504;width:536px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-femelle-accueil.webp 1692w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-femelle-accueil-768x683.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-femelle-accueil-1536x1365.webp 1536w\" data-sizes=\"(max-width: 1692px) 100vw, 1692px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Anax imperator femelle, La Croix Fourreau (France-49), 25\/08\/2019<\/figcaption><\/figure>\n\n\n\n<p><strong>Les femelles<\/strong> portent le m\u00eame pentagone sur le front, les yeux sont verts comme le thorax, l&rsquo;abdomen est vert, parfois bleu, souvent avec une ligne sombre plus ou moins large sur sa face dorsale. Lorsque l&rsquo;abdomen est bleu, elles portent les m\u00eames taches bleues avant les ailes.<br>Elles sont fr\u00e9quemment rencontr\u00e9es en ponte, seules,  alors que les femelles des autres esp\u00e8ces cit\u00e9es sont beaucoup plus discr\u00e8tes, voire rarement photographi\u00e9es. Elles s&rsquo;en distinguent par les m\u00eames crit\u00e8res de coloration thoracique que  pour les m\u00e2les. Les femelles B. pratense et A. affinis sont jaunes, les femelles A. parthenope ont les ailes color\u00e9es entre le nodus et le pt\u00e9rostigma (<a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/anax-imperator-ponte-14.jpg\">A. imperator<\/a> entre le triangle et l&rsquo;apex), les femelles A. epipphiger sont compl\u00e8tement ternes.<\/p>\n\n\n\n<h5><span style=\"text-decoration: underline;\">Habitat et distribution g\u00e9ographique<\/span><\/h5>\n\n\n\n<p>L&rsquo;Anax imperator accepte toutes sortes de milieux en France, il est cependant plus commun sur les eaux stagnantes ensoleill\u00e9es et on peut le rencontrer sur des eaux faiblement courantes. Il lui importe peu que les eaux soient acides comme en tourbi\u00e8res, saum\u00e2tres ou pollu\u00e9es, elles peuvent m\u00eame \u00eatre peu v\u00e9g\u00e9talis\u00e9es pour peu qu&rsquo;il y ait des supports d&rsquo;\u00e9mergences et des arbres \u00e0 proximit\u00e9. Il lui faut un minimum d&rsquo;espace pour d\u00e9velopper son vol et s&rsquo;il lui faut plus qu&rsquo;une simple flaque pour \u00eatre heureux, il est peu commun sur les tr\u00e8s grandes pi\u00e8ces d&rsquo;eau.<br><br>En France, il est commun et pr\u00e9sent partout, m\u00eame s&rsquo;il est beaucoup plus abondant en plaine et en dessous de 1600 m\u00e8tres (2).<br><a href=\"https:\/\/atlas-odonates.insectes.org\/odonates-de-france\/anax-imperator\">Atlas Dynamique des Odonates de France<\/a><br><br>C&rsquo;est une esp\u00e8ce Africaine qui \u00e9tend r\u00e9guli\u00e8rement et rapidement son aire de distribution gigantesque vers le nord. On le rencontre maintenant de la Su\u00e8de au sud de l&rsquo;Afrique (ici, <a href=\"https:\/\/www.meslibellules.fr\/voyages\/afrique-du-sud\/anax-imperator\/anax-imperator.php\" target=\"_blank\" rel=\"noreferrer noopener\">en Afrique du Sud<\/a>), il est pr\u00e9sent \u00e0 Madagascar et <a href=\"https:\/\/www.meslibellules.fr\/voyages\/la-reunion\/anax-imperator-mauricianus\/anax-imperator-mauricianus.php\" target=\"_blank\" rel=\"noreferrer noopener\">\u00e0 la R\u00e9union<\/a>. Il est pr\u00e9sent dans toute l&rsquo;Europe, sauf au nord du Royaume-Uni et au nord de la Su\u00e8de, et de plus en plus rare et \u00e9parpill\u00e9 jusqu&rsquo;au sud-ouest de l&rsquo;Asie (3). <br><a href=\"https:\/\/www.iucnredlist.org\/species\/59812\/72311295\">IUCN Red List<\/a><\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-male-predation-sur-le-Bombyx-du-Chene.webp\"><img decoding=\"async\" width=\"1744\" height=\"1204\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-male-predation-sur-le-Bombyx-du-Chene.webp\" alt=\"Anax imperator m\u00e2le, pr\u00e9dation sur le Bombyx du Ch\u00eane, St R\u00e9my en Mauges (F-49), 15\/08\/2015\" class=\"wp-image-24864 lazyload\" style=\"--smush-placeholder-width: 1744px; --smush-placeholder-aspect-ratio: 1744\/1204;width:540px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-male-predation-sur-le-Bombyx-du-Chene.webp 1744w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-male-predation-sur-le-Bombyx-du-Chene-768x530.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-male-predation-sur-le-Bombyx-du-Chene-1536x1060.webp 1536w\" data-sizes=\"(max-width: 1744px) 100vw, 1744px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Anax imperator m\u00e2le, pr\u00e9dation sur le Bombyx du Ch\u00eane, St R\u00e9my en Mauges (F-49), 15\/08\/2015<\/figcaption><\/figure>\n\n\n\n<h5><span style=\"text-decoration: underline;\">P\u00e9riode de vol<\/span><\/h5>\n\n\n\n<p>En France, on le signale de la mi-avril au d\u00e9but de novembre, mais il est vraiment tr\u00e8s abondant en juin et juillet.<\/p>\n\n\n\n<h5><span style=\"text-decoration: underline;\">Comportement -\u00c9cologie<\/span><\/h5>\n\n\n\n<p>Je ne peux que recommander de lire les pages \u00e9crites par Andr\u00e9 Robert (4) au sujet de cet Anax, alliant pr\u00e9cision naturaliste et po\u00e9sie, dans un d\u00e9licieux style \u00ab\u00a0\u00e0 l&rsquo;ancienne\u00a0\u00bb ; beaucoup des d\u00e9tails que je donne ici en sont issus.<br>Bien \u00e9videmment, si l&rsquo;on veut consulter une \u00e9tude scientifique sur l&rsquo;Histoire Naturelle de l&rsquo;Anax imperator, le document de 70 pages de P. Corbet (5) est incontournable et consultable sur le Web.<br><br>On l&rsquo;observe le plus souvent les effectuant des allers-retours le long de la berge de l&rsquo;\u00e9tang, chassant les intrus, poursuivant et \u00e9cartant les concurrents, pour revenir \u00e0 son incessant man\u00e8ge, qu&rsquo;il ponctue parfois de court vol stationnaire ; il fait alors les d\u00e9lices des photographes ! Mais son but est bien de d\u00e9fendre son territoire et de trouver une femelle occup\u00e9e \u00e0 pondre pour s&rsquo;accoupler.<br>Les <a href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax-imperator-accouplement-2\/anax-imperator-accouplement-5big.jpg\">accouplements<\/a> sont cependant rarement observ\u00e9s, se d\u00e9roulent \u00e0 l&rsquo;abri et durent d&rsquo;une dizaine de minutes \u00e0 une heure (2) ou cinq \u00e0 dix minutes (4). La femelle reprend alors discr\u00e8tement le chemin de la ponte pour d\u00e9poser ses \u0153ufs \u00e0 des endroits soigneusement choisis, se posant sur des champs de plantes aquatiques (potamots) ou \u00e0 d\u00e9faut des tiges de v\u00e9g\u00e9taux d\u00e9compos\u00e9s ou <a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/anax-imperator-ponte-16.jpg\">du bois mort<\/a> o\u00f9 elle ins\u00e8re ses \u0153ufs. Il s&rsquo;\u00e9coule ensuite de 15 \u00e0 50 jours entre la ponte et l&rsquo;\u00e9closion.<br>La phase larvaire dure 3 \u00e0 4 mois \u00e0 2 ans, en fonction des conditions climatiques, expliquant que l&rsquo;esp\u00e8ce soit <a href=\"https:\/\/www.meslibellules.fr\/pagesweb\/divers\/glossaire.php\">bivoltine<\/a> dans le sud de son aire, <a href=\"https:\/\/www.meslibellules.fr\/pagesweb\/divers\/glossaire.php\">univoltine<\/a> en France et<a href=\"https:\/\/www.meslibellules.fr\/pagesweb\/divers\/glossaire.php\"> semivoltine<\/a> en Europe du Nord ou en montagne.<br>Les \u00e9mergences, d&rsquo;abord massives et synchronis\u00e9es en quelques jours du printemps, peuvent laisser de <a href=\"https:\/\/www.meslibellules.fr\/blog\/2021\/05\/29\/anax-imperator-exuvies-3-3\/\" target=\"_blank\" rel=\"noreferrer noopener\">spectaculaires empilements d&rsquo;exuvies<\/a>, et vont ensuite s&rsquo;\u00e9taler sur l&rsquo;\u00e9t\u00e9 (5). Elles se produisent en fin de nuit et dure plus ou moins longtemps selon la temp\u00e9rature.<\/p>\n\n\n\n<div style=\"height:25px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<h5><span style=\"text-decoration: underline;\">\u00c9tymologie<\/span><\/h5>\n\n\n\n<div class=\"wp-block-media-text is-stacked-on-mobile\" style=\"grid-template-columns:47% auto\"><figure class=\"wp-block-media-text__media\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-ailes-dun-emergent.webp\"><img decoding=\"async\" width=\"872\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-ailes-dun-emergent.webp\" alt=\"Ailes avant expansion d'un Anax imperator m\u00e2le \u00e9mergent, St R\u00e9my en Mauges (F-49), 13\/05\/2010\" class=\"wp-image-24866 size-full lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-ailes-dun-emergent.webp 872w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Anax-imperator-ailes-dun-emergent-768x1325.webp 768w\" data-sizes=\"(max-width: 872px) 100vw, 872px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 872px; --smush-placeholder-aspect-ratio: 872\/1504;\" \/><\/a><\/figure><div class=\"wp-block-media-text__content\">\n<p>Il faut tout d&rsquo;abord revenir avec R\u00e9gis Krieg-Jacquier (6) sur la r\u00e9cente modification (2021) de la nomenclature au sujet de ce Blue emperor (comme l&rsquo;appellent les anglo-saxons), je le cite int\u00e9gralement : \u00ab\u00a0&#8230;<em>Il s\u2019agit parfois de modifications dans la mention des auteurs des descriptions originales des esp\u00e8ces comme ce fut le cas en 2021 pour Anax imperator Leach, 1815 qui devenait ainsi Anax imperator Leach in Brewster, 1815, la r\u00e9f\u00e9rence ant\u00e9rieurement utilis\u00e9e (cf. p. ex. Bridges, 1994) \u00e9tant inexacte. En effet, tous les genres attribu\u00e9s jusque-l\u00e0 \u00e0 Leach, 1815 sont maintenant \u00e0 attribuer \u00e0 Leach in Brewster, 1815. Le genre Anax est nomm\u00e9 page 137 du tome 9 de Brewster (1830), dans le chapitre Entomologie (pp. 57-172), avec une seule esp\u00e8ce cit\u00e9e (A. imperator), sur la foi d\u2019une description de Leach, et la description du genre devient donc \u00e9galement la description de cette esp\u00e8ce. Ce volume est dat\u00e9 MDCCCXXX (1830) comme tous les autres volumes (date du dernier volume de cette encyclop\u00e9die). Toutefois, le fascicule contenant la description du genre Anax a \u00e9t\u00e9 publi\u00e9 en 1815.<\/em>\u00ab\u00a0<\/p>\n\n\n\n<p><\/p>\n\n\n\n<p><span style=\"font-size: 10pt;\">Ailes avant expansion d&rsquo;un Anax imperator m\u00e2le \u00e9mergent, St R\u00e9my en Mauges (F-49), 13\/05\/2010<\/span><\/p>\n<\/div><\/div>\n\n\n\n<div style=\"height:21px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<p><strong>Anax<\/strong>, d\u2019un mot grec signifiant&nbsp;<em>roi<\/em>,&nbsp;<em>souverain<\/em>&nbsp;et est attribu\u00e9 au caract\u00e8re dominant que sugg\u00e8rent sa grande taille et son comportement (7)<br><strong>Imperator<\/strong>, du latin<em> imperator<\/em>, qui signifie <em>chef<\/em> puis <em>empereur<\/em>, insiste sur le caract\u00e8re <em>apparemment<\/em> dominant du plus commun des insectes g\u00e9ants d&rsquo;Europe.<\/p>\n\n\n\n<div style=\"height:30px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<div class=\"wp-block-buttons is-layout-flex wp-block-buttons-is-layout-flex\">\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/emerg1_1.php\">\u00c9mergence 1<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax-imperator-emergence-2-1.php\">\u00c9mergence 2<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax-imperator-incident-emergence.php\">Incident d&rsquo;\u00e9mergence<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax-imperator-exuvie-1.php\">Exuvie<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax-imperator-femelle-ailes.php\">Ailes de femelles<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax-imperator-oeil.php\">L&rsquo;\u0153il<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/femelle1.php\">Femelle<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax-imperator-femelle-immature.php\">Femelle immature<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/male1.php\">M\u00e2le<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax-imperator-accouplement-1.php\">Accouplement<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax_ponte1.php\">La Ponte<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/aeshnidae\/anax\/anax_imperator\/anax-imperator-male-en-vol.php\">M\u00e2le en vol<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/blog\/2025\/05\/21\/predation-par-anax-imperator\/\">Pr\u00e9dation sur C. aenea<\/a><\/div>\n<\/div>\n\n\n\n<div style=\"height:59px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<p>1- Leach, W. E. (1815). Entomology. In: <a href=\"https:\/\/www.biodiversitylibrary.org\/page\/17493627#page\/145\/mode\/1up\">Brewster, D. (ed.) The Edinburgh Encyclopaedia. Volume IX<\/a>. Edinburgh: William<br>Blackwood: 137<br>2- Daniel Grand et Jean-Pierre Boudot, Les Libellules de France, Belgique et Luxembourg, collection Parth\u00e9nope, 2006<br>3- <a href=\"https:\/\/www.researchgate.net\/publication\/291165298_Atlas_of_the_European_dragonflies_and_damselflies\">Atlas of the European dragonflies and damselfies<\/a>, Jean-Pierre Boudot &amp; Vincent J. Kalkman, KNNV Publishing, 2015- P. 173-175<br>4- Paul Andr\u00e9 Robert, Les Libellules, Delachaux et Niestl\u00e9, 1958<br>5- Philip S. Corbet, 1957 &#8211; <a href=\"https:\/\/www.researchgate.net\/publication\/273244224_The_Life-History_of_the_Emperor_Dragonfly_Anax_imperator_Leach_Odonata_Aeshnidae\">The Life-History of the Emperor Dragonfly Anax imperator Leach (Odonata: Aeshnidae)<\/a> &#8211; Journal of Animal Ecology 26(1):1 &#8211; May 1957<br>6- Krieg-Jacquier R., 2024 &#8211; <a href=\"https:\/\/martinia.insectes.org\/wp-content\/uploads\/2024\/07\/Krieg-Jacquier-2024-Isoaeschna-isoceles-O.F.-Muller-1767-un-nouveau.pdf\">Isoaeschna isoceles (O.F. M\u00fcller, 1767) un nouveau nom binominal pour l\u2019Aeschne isoc\u00e8le (Odonata : Aeshnidae)<\/a> &#8211; Martinia, 7 ao\u00fbt 2024<br>7- Ian Endersby &amp; Heinrich Fliedner, <a href=\"https:\/\/www.researchgate.net\/publication\/283318421_The_Naming_of_Australia's_Dragonflies\">The Naming of Australia\u2019s Dragonflies<\/a>, Busybird publishing, 2015.<\/p>\n\n\n\n<div style=\"height:15px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<hr class=\"wp-block-separator has-alpha-channel-opacity\" style=\"margin-top:0;margin-bottom:0\"\/>\n\n\n\n<div style=\"height:30px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<p>P.S. Pour \u00e9crire cette page, mes recherches sur le Web m&rsquo;ont conduit \u00e0 cet article de 2014, qui m&rsquo;a laiss\u00e9, comment l&rsquo;\u00e9crire impartialement, pour le moins dubitatif : \u00ab\u00a0<em><a href=\"https:\/\/www.researchgate.net\/publication\/263164140_Effects_of_homeopathic_Anax_imperator_on_behavioural_and_pain_models_in_mice\/citations\">Effects of homeopathic Anax imperator on behavioural and pain models in mice<\/a><\/em>\u00ab\u00a0, June 2014- Homeopathy&nbsp;104(1). Les scientifiques Turcs semblent particuli\u00e8rement attach\u00e9s \u00e0 cette \u00e9tude puisqu&rsquo;on lit dans une citation r\u00e9f\u00e9rence \u00e0 cet article :\u00a0\u00bbThe homeopathic dragonfly Anax imperator medicine is intended to treat attention deficit hyperactivity disorder (ADHD), obsessive-compulsive disorder (OCD), headache, sinusitis, and influenza. In addition, it produced hyperactivity and weight loss in naive mice and had antidepressant, anxiolytic, and analgesic effects&nbsp;(Mutlu et al., 2015). <br><br>Et moi, b\u00e9otien, qui me contentait de les photographier sans imaginer leur potentiel th\u00e9rapeutique !<\/p>\n\n\n\n<p><\/p>\n","protected":false},"excerpt":{"rendered":"<p>L&rsquo;Anax imperator (Aeshnidae) est un des plus spectaculaires Odonates d&rsquo;Europe, par son vol dominant, sa tr\u00e8s grande taille (66 \u00e0 84 mm) et l&rsquo;\u00e9l\u00e9gance de sa livr\u00e9e. Il est tr\u00e8s commun en plaine. Description &#8211; Identification Les m\u00e2les matures de l&rsquo;Anax empereur (1) ont des yeux verts ou bleu-vert et portent un pentagone sur le [&hellip;]<\/p>\n","protected":false},"author":1,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"site-sidebar-layout":"default","site-content-layout":"","ast-site-content-layout":"default","site-content-style":"default","site-sidebar-style":"default","ast-global-header-display":"","ast-banner-title-visibility":"","ast-main-header-display":"","ast-hfb-above-header-display":"","ast-hfb-below-header-display":"","ast-hfb-mobile-header-display":"","site-post-title":"","ast-breadcrumbs-content":"","ast-featured-img":"","footer-sml-layout":"","ast-disable-related-posts":"","theme-transparent-header-meta":"","adv-header-id-meta":"","stick-header-meta":"","header-above-stick-meta":"","header-main-stick-meta":"","header-below-stick-meta":"","astra-migrate-meta-layouts":"set","ast-page-background-enabled":"default","ast-page-background-meta":{"desktop":{"background-color":"var(--ast-global-color-4)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"ast-content-background-meta":{"desktop":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"footnotes":""},"class_list":["post-24840","page","type-page","status-publish","hentry"],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v27.4 - 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