{"id":6571,"date":"2020-08-02T21:01:32","date_gmt":"2020-08-02T19:01:32","guid":{"rendered":"http:\/\/www.meslibellules.fr\/blog\/?page_id=6571"},"modified":"2024-01-06T21:15:47","modified_gmt":"2024-01-06T20:15:47","slug":"ischnura-elegans-les-femelles","status":"publish","type":"page","link":"https:\/\/www.meslibellules.fr\/blog\/ischnura-elegans-les-femelles\/","title":{"rendered":"Ischnura elegans : les femelles"},"content":{"rendered":"\n<p style=\"font-size:15px\">Ischnura elegans femelle : pr\u00e9sentation<br><br>Ischnura elegans, l&rsquo;Agrion \u00e9l\u00e9gant, est tr\u00e8s commun et contrairement aux autres Coenagrionidae, on rencontre  souvent les femelles. Elles sont les seules \u00e0 pr\u00e9senter une telle diversit\u00e9 puisqu&rsquo;on compte deux formes immatures <strong>tr\u00e8s diff\u00e9rentes<\/strong> qui donnent trois formes matures.<br><br>Ces<strong> immatures<\/strong> sont\u00a0:<br>\u2014 Une forme <em>violacea<\/em>, bleu lilas, taches postoculaires bleues avec une marque (spot) bleue sur le S8 (8\u00b0 segment). <br>\u2014 Une forme <em>rufescens<\/em>, orang\u00e9e, ros\u00e9e ou rose-orang\u00e9, taches postoculaires de la m\u00eame couleur, avec un spot qui deviendra rapidement bleu sur le S8, mais <strong>qui ne pr\u00e9sente pas  de ligne noire<\/strong> sur la suture hum\u00e9rale.<\/p>\n\n\n\n<figure class=\"wp-block-gallery aligncenter has-nested-images columns-default is-cropped wp-block-gallery-1 is-layout-flex wp-block-gallery-is-layout-flex\">\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/i-elegans-violacea-1-2.jpg\"><img decoding=\"async\" width=\"1463\" height=\"760\" data-id=\"6608\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/i-elegans-violacea-1-2.jpg\" alt=\"Ischnura elegans femelle immature violacea, La Renaudi\u00e8re (France - 49), 01\/08\/2010\" class=\"wp-image-6608 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-violacea-1-2.jpg 1463w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-violacea-1-2-768x399.jpg 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-violacea-1-2-1200x623.jpg 1200w\" data-sizes=\"(max-width: 1463px) 100vw, 1463px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1463px; --smush-placeholder-aspect-ratio: 1463\/760;\" \/><\/a><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/i-elegans-rufescens-1.jpg\"><img decoding=\"async\" width=\"2019\" height=\"1004\" data-id=\"6580\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/i-elegans-rufescens-1.jpg\" alt=\"Agrion \u00e9l\u00e9gant femelle immature rufescens, Etang de P\u00e9ronne (France - 49), 04\/06\/2011\" class=\"wp-image-6580 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-rufescens-1.jpg 2019w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-rufescens-1-768x382.jpg 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-rufescens-1-1536x764.jpg 1536w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-rufescens-1-1200x597.jpg 1200w\" data-sizes=\"(max-width: 2019px) 100vw, 2019px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 2019px; --smush-placeholder-aspect-ratio: 2019\/1004;\" \/><\/a><\/figure>\n<figcaption class=\"blocks-gallery-caption wp-element-caption\">A gauche, <em>Ischnura elegans femelle immature violacea, La Renaudi\u00e8re (France &#8211; 49), 01\/08\/2010<\/em>.<br>A droite, <em>Ischnura elegans femelle immature rufescens, Etang de P\u00e9ronne (France &#8211; 49), 04\/06\/2011<\/em>.<\/figcaption><\/figure>\n\n\n\n\n\n<p style=\"font-size:15px\">La forme <span class=\"underline\" style=\"text-decoration: underline\">I. e. <em>violacea<\/em><\/span> va \u00e9voluer en 2 formes <strong>matures\u00a0:<\/strong><br>\u2014 Une forme dite <em>androchrome<\/em> (1), (2), de la m\u00eame couleur que les m\u00e2les matures, bleue, avec un spot bleu sur S8. (3)<br>\u2014 Une forme dite<em> infuscans<\/em> (4), verte, taches postoculaires de la m\u00eame couleur, avec un spot sur S8  vert, qui deviendra terne ou brun verd\u00e2tre.<br><br>La forme <span class=\"underline\" style=\"text-decoration: underline\">I. e. <em>rufescens<\/em><\/span> \u00e9volue vers une forme <strong>mature<\/strong> <em>rufescens obsoleta<\/em> (5), plus sombre, terne, avec une tache sur S8 qui s&rsquo;att\u00e9nue, s&rsquo;assombrit sans g\u00e9n\u00e9ralement dispara\u00eetre. On la reconna\u00eet cependant toujours facilement \u00e0 l&rsquo;absence d&rsquo;un trait ou d&rsquo;une bande noire bien nette sur la suture hum\u00e9rale.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-large is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ischnura-elegans-matures.jpg\"><img decoding=\"async\" width=\"2042\" height=\"2368\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ischnura-elegans-matures.jpg\" alt=\"Les 3 formes de femelles matures Ischnura elegans\" class=\"wp-image-6624 lazyload\" style=\"--smush-placeholder-width: 2042px; --smush-placeholder-aspect-ratio: 2042\/2368;width:516px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures.jpg 2042w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures-768x891.jpg 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures-1325x1536.jpg 1325w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures-1766x2048.jpg 1766w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures-1200x1392.jpg 1200w\" data-sizes=\"(max-width: 2042px) 100vw, 2042px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Les 3 formes de femelles matures Ischnura elegans. En haut I. e. androchrome, au milieu I. e. infuscans, en bas I. e. rufescens obsoleta.<\/figcaption><\/figure>\n\n\n\n<p style=\"font-size:15px\">Tout ceci peut para\u00eetre un peu confus ou en tout cas complexe et on trouve un sch\u00e9ma tr\u00e8s int\u00e9ressant  qui r\u00e9sume tout ceci dans<em> Polyandry and alternative mating tactics<\/em>, Bryan D. Neff and Erik I. Svensson, Phil. Trans. R. Soc. B 2013 368, 20120045, published online 21 January 2013 (sur ce document, j&rsquo;ai modifi\u00e9 infuscans obsoleta en rufescens obsoleta).<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-large is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ontogenie-fem-ischneleg-1.jpg\"><img decoding=\"async\" width=\"1209\" height=\"760\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ontogenie-fem-ischneleg-1.jpg\" alt=\"Ontog\u00e9nie du d\u00e9veloppement de la couleur pour Ischnura elegans femelle.\" class=\"wp-image-6634 lazyload\" style=\"--smush-placeholder-width: 1209px; --smush-placeholder-aspect-ratio: 1209\/760;width:544px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ontogenie-fem-ischneleg-1.jpg 1209w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ontogenie-fem-ischneleg-1-768x483.jpg 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ontogenie-fem-ischneleg-1-1200x754.jpg 1200w\" data-sizes=\"(max-width: 1209px) 100vw, 1209px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Ontog\u00e9nie du d\u00e9veloppement de la couleur pour Ischnura elegans femelle.<\/figcaption><\/figure>\n\n\n\n<p style=\"font-size:15px\">Cette volont\u00e9 de ranger chaque femelle dans une bo\u00eete diff\u00e9rente, mature\/immature par exemple, se heurte \u00e0 la r\u00e9alit\u00e9 du terrain, puisqu&rsquo;elles ne deviennent pas matures <em>tout d&rsquo;un coup<\/em>. On peut donc trouver tous les interm\u00e9diaires entre les formes matures et immatures, c&rsquo;est-\u00e0-dire des variantes des 5 formes de femelles. On trouve m\u00eame des femelles difficiles \u00e0 classer m\u00eame si on conna\u00eet bien toutes les formes.<br><br>Par souci de simplification, K.-D. B. Dijkstra, dans <em>Guide des Libellules de France et d&rsquo;Europe<\/em>, 2007, a regroup\u00e9 les formes immatures et matures afin de cr\u00e9er les types A, B et C, comme on le voit ci-dessous. Ne nous voilons pas la face, c&rsquo;est tr\u00e8s commode, car comme dit plus haut, d\u00e9finir si une  femelle est mature ou immature est parfois tr\u00e8s d\u00e9licat et c&rsquo;est ce que j&rsquo;ai adopt\u00e9 dans les pages suivantes, m\u00eame si je me suis risqu\u00e9 souvent \u00e0 s\u00e9parer les immatures et les matures.<\/p>\n\n\n\n<p style=\"font-size:16px\">Les femelles des formes <strong><em>violacea<\/em> et <em>androchrome<\/em><\/strong> (type A) <a href=\"http:\/\/www.meslibellules.fr\/pagesweb\/coenagrionidae\/ischnura\/ischnura-elegans\/femmella1.php\"><strong>\u2192<\/strong><\/a><br>Les femelles <em><strong>infuscans<\/strong><\/em>  (type B)  <a href=\"http:\/\/www.meslibellules.fr\/pagesweb\/coenagrionidae\/ischnura\/ischnura-elegans\/femelleb1.php\">\u2192<\/a><br>Les femelles <strong><em>rufescens<\/em> et <em>rufescens obsoleta<\/em><\/strong>  (type C) <a href=\"http:\/\/www.meslibellules.fr\/pagesweb\/coenagrionidae\/ischnura\/ischnura-elegans\/femellec1.php\"> \u2192<\/a><\/p>\n\n\n\n<div style=\"height:38px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<figure class=\"wp-block-image size-large is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/ischnura-elegans-3\/\"><img decoding=\"async\" width=\"100\" height=\"20\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/2019\/05\/accueilielegans.gif\" alt=\"\" class=\"wp-image-2893 lazyload\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 100px; --smush-placeholder-aspect-ratio: 100\/20;\" \/><\/a><\/figure>\n\n\n\n<div style=\"height:55px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<p class=\"has-small-font-size\">-1- Androchrome signifie <em>de la m\u00eame couleur que le m\u00e2le<\/em>.<br>-2- Parfois appel\u00e9e <em>typica<\/em><br>-3-De tr\u00e8s rares formes androchromes ne passent pas par le stade violacea mais par une forme vert clair comme les m\u00e2les immatures&#8230; voir <em>Reflectance spectra and mating patterns support intraspecific mimicry in the colour polymorphic damselfly Ischnura elegans<\/em>, Hans Van Gossum \u2022 Jessica Bots \u2022 Jasper Van Heusden \u2022 Martijn Hammers \u2022 Katleen Huyghe \u2022 Nathan I. Morehouse, Evol Ecol (2011) 25:139\u2013154<br>-4- Infuscans du latin <em>infusco<\/em>, signifie <em>devenir brun, devenir terne<\/em>.<br>-5- &#8230; parfois appel\u00e9, de fa\u00e7on incompr\u00e9hensible pour moi, infuscans obsoleta , voir <em>The Dragonglies of Europe<\/em>, Askew 2004, ou d&rsquo;autres publication scientifiques. Mais dans Michael J. Parr&nbsp;(1999)&nbsp;THE TERMINOLOGY OF FEMALE POLYMORPHS OF&nbsp;<em>ISCHNURA<\/em>&nbsp;(ZYGOPTERA: COENAGRIONIDAE),&nbsp;International Journal of Odonatology,&nbsp;2:1,&nbsp;95-99,&nbsp;DOI:&nbsp;<a href=\"https:\/\/doi.org\/10.1080\/13887890.1999.9748116\">10.1080\/13887890.1999.9748116<\/a>  qui n&rsquo;a \u00e9t\u00e9 mis enligne qu&rsquo;en 2012, il est logiquement sugg\u00e9r\u00e9 que cette forme soit appel\u00e9 <em>rufescens obsoleta<\/em>.<\/p>\n","protected":false},"excerpt":{"rendered":"<p>Ischnura elegans femelle : pr\u00e9sentation Ischnura elegans, l&rsquo;Agrion \u00e9l\u00e9gant, est tr\u00e8s commun et contrairement aux autres Coenagrionidae, on rencontre souvent les femelles. Elles sont les seules \u00e0 pr\u00e9senter une telle diversit\u00e9 puisqu&rsquo;on compte deux formes immatures tr\u00e8s diff\u00e9rentes qui donnent trois formes matures. Ces immatures sont\u00a0:\u2014 Une forme violacea, bleu lilas, taches postoculaires bleues avec [&hellip;]<\/p>\n","protected":false},"author":1,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"open","ping_status":"closed","template":"","meta":{"site-sidebar-layout":"default","site-content-layout":"","ast-site-content-layout":"default","site-content-style":"default","site-sidebar-style":"default","ast-global-header-display":"","ast-banner-title-visibility":"","ast-main-header-display":"","ast-hfb-above-header-display":"","ast-hfb-below-header-display":"","ast-hfb-mobile-header-display":"","site-post-title":"","ast-breadcrumbs-content":"","ast-featured-img":"","footer-sml-layout":"","ast-disable-related-posts":"","theme-transparent-header-meta":"","adv-header-id-meta":"","stick-header-meta":"","header-above-stick-meta":"","header-main-stick-meta":"","header-below-stick-meta":"","astra-migrate-meta-layouts":"set","ast-page-background-enabled":"default","ast-page-background-meta":{"desktop":{"background-color":"var(--ast-global-color-4)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"ast-content-background-meta":{"desktop":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"footnotes":""},"class_list":["post-6571","page","type-page","status-publish","hentry"],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v27.3 - 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