{"id":6571,"date":"2020-08-02T21:01:32","date_gmt":"2020-08-02T19:01:32","guid":{"rendered":"http:\/\/www.meslibellules.fr\/blog\/?page_id=6571"},"modified":"2024-01-06T21:15:47","modified_gmt":"2024-01-06T20:15:47","slug":"ischnura-elegans-les-femelles","status":"publish","type":"page","link":"https:\/\/www.meslibellules.fr\/blog\/ischnura-elegans-les-femelles\/","title":{"rendered":"Ischnura elegans : les femelles"},"content":{"rendered":"\n<p class=\"wp-block-paragraph\" style=\"font-size:15px\">Ischnura elegans femelle : pr\u00e9sentation<br><br>Ischnura elegans, l'Agrion \u00e9l\u00e9gant, est tr\u00e8s commun et contrairement aux autres Coenagrionidae, on rencontre  souvent les femelles. Elles sont les seules \u00e0 pr\u00e9senter une telle diversit\u00e9 puisqu'on compte deux formes immatures <strong>tr\u00e8s diff\u00e9rentes<\/strong> qui donnent trois formes matures.<br><br>Ces<strong> immatures<\/strong> sont\u00a0:<br>\u2014 Une forme <em>violacea<\/em>, bleu lilas, taches postoculaires bleues avec une marque (spot) bleue sur le S8 (8\u00b0 segment). <br>\u2014 Une forme <em>rufescens<\/em>, orang\u00e9e, ros\u00e9e ou rose-orang\u00e9, taches postoculaires de la m\u00eame couleur, avec un spot qui deviendra rapidement bleu sur le S8, mais <strong>qui ne pr\u00e9sente pas  de ligne noire<\/strong> sur la suture hum\u00e9rale.<\/p>\n\n\n\n<figure class=\"wp-block-gallery aligncenter has-nested-images columns-default is-cropped wp-block-gallery-1 is-layout-flex wp-block-gallery-is-layout-flex\">\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/i-elegans-violacea-1-2.jpg\"><img decoding=\"async\" width=\"1463\" height=\"760\" data-id=\"6608\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/i-elegans-violacea-1-2.jpg\" alt=\"Ischnura elegans femelle immature violacea, La Renaudi\u00e8re (France - 49), 01\/08\/2010\" class=\"wp-image-6608 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-violacea-1-2.jpg 1463w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-violacea-1-2-768x399.jpg 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-violacea-1-2-1200x623.jpg 1200w\" data-sizes=\"(max-width: 1463px) 100vw, 1463px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1463px; --smush-placeholder-aspect-ratio: 1463\/760;\" \/><\/a><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/i-elegans-rufescens-1.jpg\"><img decoding=\"async\" width=\"2019\" height=\"1004\" data-id=\"6580\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/i-elegans-rufescens-1.jpg\" alt=\"Agrion \u00e9l\u00e9gant femelle immature rufescens, Etang de P\u00e9ronne (France - 49), 04\/06\/2011\" class=\"wp-image-6580 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-rufescens-1.jpg 2019w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-rufescens-1-768x382.jpg 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-rufescens-1-1536x764.jpg 1536w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/i-elegans-rufescens-1-1200x597.jpg 1200w\" data-sizes=\"(max-width: 2019px) 100vw, 2019px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 2019px; --smush-placeholder-aspect-ratio: 2019\/1004;\" \/><\/a><\/figure>\n<figcaption class=\"blocks-gallery-caption wp-element-caption\">A gauche, <em>Ischnura elegans femelle immature violacea, La Renaudi\u00e8re (France - 49), 01\/08\/2010<\/em>.<br>A droite, <em>Ischnura elegans femelle immature rufescens, Etang de P\u00e9ronne (France - 49), 04\/06\/2011<\/em>.<\/figcaption><\/figure>\n\n\n\n\n\n<p class=\"wp-block-paragraph\" style=\"font-size:15px\">La forme <span class=\"underline\" style=\"text-decoration: underline\">I. e. <em>violacea<\/em><\/span> va \u00e9voluer en 2 formes <strong>matures\u00a0:<\/strong><br>\u2014 Une forme dite <em>androchrome<\/em> (1), (2), de la m\u00eame couleur que les m\u00e2les matures, bleue, avec un spot bleu sur S8. (3)<br>\u2014 Une forme dite<em> infuscans<\/em> (4), verte, taches postoculaires de la m\u00eame couleur, avec un spot sur S8  vert, qui deviendra terne ou brun verd\u00e2tre.<br><br>La forme <span class=\"underline\" style=\"text-decoration: underline\">I. e. <em>rufescens<\/em><\/span> \u00e9volue vers une forme <strong>mature<\/strong> <em>rufescens obsoleta<\/em> (5), plus sombre, terne, avec une tache sur S8 qui s'att\u00e9nue, s'assombrit sans g\u00e9n\u00e9ralement dispara\u00eetre. On la reconna\u00eet cependant toujours facilement \u00e0 l'absence d'un trait ou d'une bande noire bien nette sur la suture hum\u00e9rale.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-large is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ischnura-elegans-matures.jpg\"><img decoding=\"async\" width=\"2042\" height=\"2368\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ischnura-elegans-matures.jpg\" alt=\"Les 3 formes de femelles matures Ischnura elegans\" class=\"wp-image-6624 lazyload\" style=\"--smush-placeholder-width: 2042px; --smush-placeholder-aspect-ratio: 2042\/2368;width:516px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures.jpg 2042w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures-768x891.jpg 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures-1325x1536.jpg 1325w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures-1766x2048.jpg 1766w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ischnura-elegans-matures-1200x1392.jpg 1200w\" data-sizes=\"(max-width: 2042px) 100vw, 2042px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Les 3 formes de femelles matures Ischnura elegans. En haut I. e. androchrome, au milieu I. e. infuscans, en bas I. e. rufescens obsoleta.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\" style=\"font-size:15px\">Tout ceci peut para\u00eetre un peu confus ou en tout cas complexe et on trouve un sch\u00e9ma tr\u00e8s int\u00e9ressant  qui r\u00e9sume tout ceci dans<em> Polyandry and alternative mating tactics<\/em>, Bryan D. Neff and Erik I. Svensson, Phil. Trans. R. Soc. B 2013 368, 20120045, published online 21 January 2013 (sur ce document, j'ai modifi\u00e9 infuscans obsoleta en rufescens obsoleta).<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-large is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ontogenie-fem-ischneleg-1.jpg\"><img decoding=\"async\" width=\"1209\" height=\"760\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ontogenie-fem-ischneleg-1.jpg\" alt=\"Ontog\u00e9nie du d\u00e9veloppement de la couleur pour Ischnura elegans femelle.\" class=\"wp-image-6634 lazyload\" style=\"--smush-placeholder-width: 1209px; --smush-placeholder-aspect-ratio: 1209\/760;width:544px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ontogenie-fem-ischneleg-1.jpg 1209w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ontogenie-fem-ischneleg-1-768x483.jpg 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/\/ontogenie-fem-ischneleg-1-1200x754.jpg 1200w\" data-sizes=\"(max-width: 1209px) 100vw, 1209px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Ontog\u00e9nie du d\u00e9veloppement de la couleur pour Ischnura elegans femelle.<\/figcaption><\/figure>\n\n\n\n<p class=\"wp-block-paragraph\" style=\"font-size:15px\">Cette volont\u00e9 de ranger chaque femelle dans une bo\u00eete diff\u00e9rente, mature\/immature par exemple, se heurte \u00e0 la r\u00e9alit\u00e9 du terrain, puisqu'elles ne deviennent pas matures <em>tout d'un coup<\/em>. On peut donc trouver tous les interm\u00e9diaires entre les formes matures et immatures, c'est-\u00e0-dire des variantes des 5 formes de femelles. On trouve m\u00eame des femelles difficiles \u00e0 classer m\u00eame si on conna\u00eet bien toutes les formes.<br><br>Par souci de simplification, K.-D. B. Dijkstra, dans <em>Guide des Libellules de France et d'Europe<\/em>, 2007, a regroup\u00e9 les formes immatures et matures afin de cr\u00e9er les types A, B et C, comme on le voit ci-dessous. Ne nous voilons pas la face, c'est tr\u00e8s commode, car comme dit plus haut, d\u00e9finir si une  femelle est mature ou immature est parfois tr\u00e8s d\u00e9licat et c'est ce que j'ai adopt\u00e9 dans les pages suivantes, m\u00eame si je me suis risqu\u00e9 souvent \u00e0 s\u00e9parer les immatures et les matures.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\" style=\"font-size:16px\">Les femelles des formes <strong><em>violacea<\/em> et <em>androchrome<\/em><\/strong> (type A) <a href=\"http:\/\/www.meslibellules.fr\/pagesweb\/coenagrionidae\/ischnura\/ischnura-elegans\/femmella1.php\"><strong>\u2192<\/strong><\/a><br>Les femelles <em><strong>infuscans<\/strong><\/em>  (type B)  <a href=\"http:\/\/www.meslibellules.fr\/pagesweb\/coenagrionidae\/ischnura\/ischnura-elegans\/femelleb1.php\">\u2192<\/a><br>Les femelles <strong><em>rufescens<\/em> et <em>rufescens obsoleta<\/em><\/strong>  (type C) <a href=\"http:\/\/www.meslibellules.fr\/pagesweb\/coenagrionidae\/ischnura\/ischnura-elegans\/femellec1.php\"> \u2192<\/a><\/p>\n\n\n\n<div style=\"height:38px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<figure class=\"wp-block-image size-large is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/ischnura-elegans-3\/\"><img decoding=\"async\" width=\"100\" height=\"20\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/2019\/05\/accueilielegans.gif\" alt=\"\" class=\"wp-image-2893 lazyload\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 100px; --smush-placeholder-aspect-ratio: 100\/20;\" \/><\/a><\/figure>\n\n\n\n<div style=\"height:55px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<p class=\"has-small-font-size wp-block-paragraph\">-1- Androchrome signifie <em>de la m\u00eame couleur que le m\u00e2le<\/em>.<br>-2- Parfois appel\u00e9e <em>typica<\/em><br>-3-De tr\u00e8s rares formes androchromes ne passent pas par le stade violacea mais par une forme vert clair comme les m\u00e2les immatures... voir <em>Reflectance spectra and mating patterns support intraspecific mimicry in the colour polymorphic damselfly Ischnura elegans<\/em>, Hans Van Gossum \u2022 Jessica Bots \u2022 Jasper Van Heusden \u2022 Martijn Hammers \u2022 Katleen Huyghe \u2022 Nathan I. Morehouse, Evol Ecol (2011) 25:139\u2013154<br>-4- Infuscans du latin <em>infusco<\/em>, signifie <em>devenir brun, devenir terne<\/em>.<br>-5- ... parfois appel\u00e9, de fa\u00e7on incompr\u00e9hensible pour moi, infuscans obsoleta , voir <em>The Dragonglies of Europe<\/em>, Askew 2004, ou d'autres publication scientifiques. Mais dans Michael J. Parr&nbsp;(1999)&nbsp;THE TERMINOLOGY OF FEMALE POLYMORPHS OF&nbsp;<em>ISCHNURA<\/em>&nbsp;(ZYGOPTERA: COENAGRIONIDAE),&nbsp;International Journal of Odonatology,&nbsp;2:1,&nbsp;95-99,&nbsp;DOI:&nbsp;<a href=\"https:\/\/doi.org\/10.1080\/13887890.1999.9748116\">10.1080\/13887890.1999.9748116<\/a>  qui n'a \u00e9t\u00e9 mis enligne qu'en 2012, il est logiquement sugg\u00e9r\u00e9 que cette forme soit appel\u00e9 <em>rufescens obsoleta<\/em>.<\/p>\n","protected":false},"excerpt":{"rendered":"<p>Ischnura elegans femelle : pr\u00e9sentation Ischnura elegans, l'Agrion \u00e9l\u00e9gant, est tr\u00e8s commun et contrairement aux autres Coenagrionidae, on rencontre souvent les femelles. Elles sont les seules \u00e0 pr\u00e9senter une telle diversit\u00e9 puisqu'on compte deux formes immatures tr\u00e8s diff\u00e9rentes qui donnent trois formes matures. Ces immatures sont\u00a0:\u2014 Une forme violacea, bleu lilas, taches postoculaires bleues avec [&hellip;]<\/p>\n","protected":false},"author":1,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"open","ping_status":"closed","template":"","meta":{"site-sidebar-layout":"default","site-content-layout":"","ast-site-content-layout":"default","site-content-style":"default","site-sidebar-style":"default","ast-global-header-display":"","ast-banner-title-visibility":"","ast-main-header-display":"","ast-hfb-above-header-display":"","ast-hfb-below-header-display":"","ast-hfb-mobile-header-display":"","site-post-title":"","ast-breadcrumbs-content":"","ast-featured-img":"","footer-sml-layout":"","ast-disable-related-posts":"","theme-transparent-header-meta":"","adv-header-id-meta":"","stick-header-meta":"","header-above-stick-meta":"","header-main-stick-meta":"","header-below-stick-meta":"","astra-migrate-meta-layouts":"set","ast-page-background-enabled":"default","ast-page-background-meta":{"desktop":{"background-color":"var(--ast-global-color-4)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"ast-content-background-meta":{"desktop":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"footnotes":""},"class_list":["post-6571","page","type-page","status-publish","hentry"],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v28.1 - 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