{"id":25608,"date":"2025-07-25T08:58:23","date_gmt":"2025-07-25T06:58:23","guid":{"rendered":"https:\/\/www.meslibellules.fr\/blog\/?p=25608"},"modified":"2025-10-25T21:53:39","modified_gmt":"2025-10-25T19:53:39","slug":"neurothemis-ramburii","status":"publish","type":"post","link":"https:\/\/www.meslibellules.fr\/blog\/2025\/07\/25\/neurothemis-ramburii\/","title":{"rendered":"Born\u00e9o, Sarawak – Neurothemis ramburii (Kaup in Brauer, 1866)"},"content":{"rendered":"\n
\"Neurothemis<\/a>
Neurothemis ramburii m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025<\/figcaption><\/figure>\n\n\n\n

Je pensais avoir d\u00e9j\u00e0 rencontr\u00e9 Neurothemis ramburii en 2013, en Malaisie, identifications confirm\u00e9es par A.G. Orr (5), mais apr\u00e8s avoir progress\u00e9 dans l’identification de ces Libellulidae, notamment gr\u00e2ce \u00e0 Seehausen & Dow, 2016, il s’agirait d’une erreur et cette esp\u00e8ce ne serait pas pr\u00e9sente sur la P\u00e9ninsule (1).
\u00c0 Born\u00e9o, il n’y a que 3 Neurothemis et leur coloration alaire fait qu’il est impossible de les confondre avec un autre genre : Neurothemis ramburii, fluctuans et terminata. Sur photo, l’identification repose sur les marques noires abdominales, la forme de la zone hyaline des ailes et la nervation, en particulier du triangle de l’aile ant\u00e9rieure.
La description de l’esp\u00e8ce figure dans un ouvrage de Brauer, 1866 (2) sous le nom de Polyneura ramburii. En 1867, il cr\u00e9era le genre Neurothemis.<\/p>\n\n\n\n

\"Neurothemis<\/a>
Neurothemis ramburii m\u00e2le, Sarawak, Mount Matang, 23\/03\/2025<\/figcaption><\/figure>\n\n\n\n

On note en effet que les marques abdominales lat\u00e9rales de ces sujets touchent la partie proximale de chaque segment ou sont d\u00e9cal\u00e9es vers l’avant ; pour N. fluctuans, elles sont au milieu des segments ; pour N. terminata, elles touchent ou sont proches de l’apex des segments. <\/p>\n\n\n\n

\n
\"Neurothemis<\/a><\/figure>\n\n\n\n
\"Abdomens<\/a><\/figure>\n
Neurothemis ramburii m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025
Malte Seehausen & Rory A. Dow, 2016 (1)
<\/figcaption><\/figure>\n\n\n\n

La coloration de l’aile ant\u00e9rieure parvient ici jusqu’au d\u00e9but du nodus et rejoint le bord post\u00e9rieur de l’aile presque en ligne droite, alors que pour l’aile post\u00e9rieure, elle dessine une courbe, tr\u00e8s l\u00e9g\u00e8rement cass\u00e9e en son milieu. Pour N. fluctuans cette courbe n’est pas cass\u00e9e et se prolonge plus loin vers la base de l’aile ; pour N. terminata la coloration rejoint le bord post\u00e9rieur en ligne droite.
Noter, toujours issu du m\u00eame document, la tr\u00e8s grande variabilit\u00e9 de l’\u00e9tendue de la coloration qui a permis de d\u00e9finir plusieurs sous-esp\u00e8ces g\u00e9ographiquement localis\u00e9es.<\/p>\n\n\n\n

\"Ailes
Ailes des Neurothemis ramburii (1)<\/figcaption><\/figure>\n\n\n\n

Noter \u00e9galement la variabilit\u00e9 de l’intensit\u00e9 des marques noires sur l’abdomen, d’un sujet \u00e0 l’autre, sans doute en raison du vieillissement. La \u00ab\u00a0cassure\u00a0\u00bb de la courbe de la coloration de l’aile post\u00e9rieure est bien visible sur les sujets ci-dessous.<\/p>\n\n\n\n

\n
\"Neurothemis<\/a><\/figure>\n\n\n\n
\"Neurothemis<\/figure>\n
Neurothemis ramburii m\u00e2le, Sarawak, Matang Wildlife Center, 19\/03\/2025<\/figcaption><\/figure>\n\n\n\n

Dans Dragonflies of Japan (3) on trouve le dessin ci-dessous qui est tr\u00e8s int\u00e9ressant pour les femelles, car les triangles des ailes ant\u00e9rieures sont tr\u00e8s diff\u00e9rents. Malheureusement, je n’ai pas vu et en tout cas pas photographi\u00e9 de femelle N. ramburii. Celles-ci pr\u00e9sentent deux formes, l’une aux ailes color\u00e9es, l’autre aux ailes hyalines cependant tach\u00e9es aux apex.
En ce qui concerne les m\u00e2les, Le triangle de N. fluctuans est tr\u00e8s diff\u00e9rent des deux autres qui sont assez proches. Il ne semble pas que l’analyse des triangles apportent des \u00e9l\u00e9ments pour s\u00e9parer N. ramburii et N. terminata.<\/p>\n\n\n\n

\"Triangle<\/a>
Triangle alaire des Neurothemis fluctuans, ramburii et terminata (3)<\/figcaption><\/figure>\n\n\n\n

Il est certain qu’il faut rester prudent dans les identifications de Neurothemis et qu’il faut toujours additionner les crit\u00e8res, en particulier associer marques abdominales et coloration alaire. De plus, l’existence d’hybrides est envisag\u00e9e, entre N. ramburii et terminata et entre ce dernier et N. fluctuans (1) …

Brauer lui donne une longueur totale de 41 mm, il varie sans doute de 37 \u00e0 41 mm. Il est plus petit que N. terminata (environ 43 mm) mais nettement plus grand que N. fluctuans qui toise autour de 30 mm.<\/p>\n\n\n\n

\n
\"Neurothemis<\/a><\/figure>\n\n\n\n
\"Neurothemis<\/a><\/figure>\n
Neurothemis ramburii m\u00e2le, Sarawak, Bung Ngiraja, 26\/03\/2025<\/figcaption><\/figure>\n\n\n\n

On le trouve dans une grande diversit\u00e9 de biotopes : zones mar\u00e9cageuses en bord de for\u00eat, mares, \u00e9tangs, foss\u00e9s, rizi\u00e8res ou canaux d’irrigation. Watanabe & al. <\/em>(4) qui ont produit un document sur la larve de Neurothemis ramburii l’ont trouv\u00e9 particuli\u00e8rement abondant sur les foss\u00e9s agricoles, surtout l\u00e0 o\u00f9 il y avait quelques centim\u00e8tres de sable.
Ils d\u00e9crivent que la copulation se fait en vol ou les sujets pos\u00e9s de fa\u00e7on intermittente, la ponte s’ensuit avec la femelle qui frappe l’eau de l’extr\u00e9mit\u00e9 de son abdomen, gard\u00e9e par le m\u00e2le.
Je suis tout de m\u00eame surpris que la photo qui illustre cet article montre des caract\u00e8res alaires qui me paraissent incompatibles avec ce que l’on lit dans Seehausen & Dow, 2016 (1).

D’ailleurs, si l’on suit les m\u00eames auteurs, la distribution de l’esp\u00e8ce n’atteint pas ces iles \u00e0 l’est de Ta\u00efwan.
On le rencontre \u00e0 Born\u00e9o, Java, dans les Petites Iles de la Sonde, aux Moluques, dans les iles de la c\u00f4te ouest de la Nouvelle-Guin\u00e9e, aux Philippines, \u00e0 Sumatra et au Sulawesi.
IUCN Red List<\/a>.<\/p>\n\n\n\n

\n
\"Neurothemis<\/a><\/figure>\n\n\n\n
\"Neurothemis<\/a>
<\/figcaption><\/figure>\n
Neurothemis ramburii m\u00e2le, Sarawak, Matang Wildlife Center, 19\/03\/2025 & Bung Ngiraja, 26\/03\/2025<\/figcaption><\/figure>\n\n\n\n

Quelques mois apr\u00e8s avoir r\u00e9dig\u00e9 cette page sur les m\u00e2les, j’ai d\u00e9couvert quelques photos d’une femelle prise dans une zone de for\u00eat assez dense o\u00f9 nous n’avons pas rencontr\u00e9 de m\u00e2les.
Son identification repose sur la lame vulvaire saillante, longue et orthogonale, et le motif \u00ab\u00a0en \u00e9chelle\u00a0\u00bb tr\u00e8s gras de la face dorsale de son abdomen, la pr\u00e9sence d’une seule veine transverse dans l’espace cubito-anal, comme il est montr\u00e9 dans le document (1).<\/p>\n\n\n\n

\n
\"Neurothemis<\/a><\/figure>\n\n\n\n
\"Neurothemis<\/a><\/figure>\n
Neurothemis ramburii femelle, Sarawak, Ranchan Recreational Park, 30\/03\/2025<\/figcaption><\/figure>\n\n\n\n
<\/div>\n\n\n\n

\u00c9tymologie<\/strong>
Neurothemis<\/strong> (6), du grec neuro<\/em> pour \u2013nerf <\/em>ou plut\u00f4t \u2013nervure des ailes<\/em> en entomologie, et de Themis \u2013 d\u00e9esse grecque de l\u2019ordre et de la justice, qui a \u00e9t\u00e9 utilis\u00e9 de nombreuses fois depuis Hagen (1861). Le nom de la d\u00e9esse de l\u2019ordre est particuli\u00e8rement bien adapt\u00e9 \u00e0 la taxonomie, science qui vise \u00e0 d\u00e9crire et ordonner les familles, genres et esp\u00e8ces. Brauer a cr\u00e9\u00e9 ce nom de genre en 1867 pour remplacer le genre Polyneura, cr\u00e9\u00e9 par Rambur en 1842, car ce dernier \u00e9tait d\u00e9j\u00e0 utilis\u00e9 chez les H\u00e9mipt\u00e8res. Ce nom de genre, comme le nom actuel, \u00e9tait destin\u00e9 \u00e0 insister sur le grand nombre de cellules de leurs ailes.
Ramburii<\/strong> est bien s\u00fbr un hommage \u00e0 Jules Rambur, m\u00e9decin et entomologiste, descripteur de nombreuses esp\u00e8ces d’Odonates et r\u00e9dacteur de la fameuse Histoire Naturelle des Insectes (1842).<\/p>\n\n\n\n

<\/div>\n\n\n\n

1- Malte Seehausen & Rory A. Dow, 2016 – Morphological studies and taxonomic considerations on the \u2018reddishbrownwinged\u2019 group of Neurothemis Brauer, 1867 with the description of N. taiwanensis sp. nov. (Odonata: Libellulidae)<\/a> – Journal of the International Dragonfly Fund.
2- Brauer, F. 1866. Beschreibung neuer exotischer Libellen<\/a>. Verhandlungen der Kaiserlich-K\u00f6niglichen Zoologisch-Botanischen Gesellschaft Wien 16: 563-570. P. 568.
3- Akira Ozono, Itsura Kawashima, Ryo Futahashi, 2012 – Dragonflies of Japan – Bun-ichi Sogo Shuppan Co.
4- Ken\u2019ichi Watanabe, Itsuro Kawashima & Akihiko Sasamoto, 2013 – Notes on the larva of Neurothemis ramburii ramburii (Kaup in Brauer, 1866) obtained from Iriomote-jima Island, Yaeyama Islands,southern Ryukyus, Japan (Anisoptera: Libellulidae)<\/a> – TOMBO, Fukui, 55: 83-87. May. 15, 2013
5- A.G. Orr, 2005. Dragonflies of Peninsular Malaysia and Singapore- Natural History Publication (Born\u00e9o) A.G.
6- The Naming of Australia\u2019s Dragonflies<\/a>, Ian Endersby & Heinrich Fliedner, Busybird publishing.<\/p>\n\n\n\n

<\/div>\n\n\n\n
\n
Accueil Sarawak<\/a><\/div>\n\n\n\n
Listes des Odonates d’Asie<\/a><\/div>\n\n\n\n
Listes continentales<\/a><\/div>\n<\/div>\n\n\n\n

<\/p>\n","protected":false},"excerpt":{"rendered":"

Je pensais avoir d\u00e9j\u00e0 rencontr\u00e9 Neurothemis ramburii en 2013, en Malaisie, identifications confirm\u00e9es par A.G. Orr (5), mais apr\u00e8s avoir progress\u00e9 dans l’identification de ces Libellulidae, notamment gr\u00e2ce \u00e0 Seehausen & Dow, 2016, il s’agirait d’une erreur et cette esp\u00e8ce ne serait pas pr\u00e9sente sur la P\u00e9ninsule (1).\u00c0 Born\u00e9o, il n’y a que 3 Neurothemis […]<\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"image","meta":{"site-sidebar-layout":"default","site-content-layout":"","ast-site-content-layout":"default","site-content-style":"default","site-sidebar-style":"default","ast-global-header-display":"","ast-banner-title-visibility":"","ast-main-header-display":"","ast-hfb-above-header-display":"","ast-hfb-below-header-display":"","ast-hfb-mobile-header-display":"","site-post-title":"","ast-breadcrumbs-content":"","ast-featured-img":"","footer-sml-layout":"","ast-disable-related-posts":"","theme-transparent-header-meta":"","adv-header-id-meta":"","stick-header-meta":"","header-above-stick-meta":"","header-main-stick-meta":"","header-below-stick-meta":"","astra-migrate-meta-layouts":"set","ast-page-background-enabled":"default","ast-page-background-meta":{"desktop":{"background-color":"var(--ast-global-color-4)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"ast-content-background-meta":{"desktop":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"footnotes":""},"categories":[38596,38606],"tags":[38599,38617],"class_list":["post-25608","post","type-post","status-publish","format-image","hentry","category-borneo-sarawak","category-libellulidae-borneo-sarawak","tag-borneo-sarawak","tag-libellulidae-borneo-2","post_format-post-format-image"],"yoast_head":"\nNeurothemis ramburii - 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