{"id":25647,"date":"2025-07-29T18:38:18","date_gmt":"2025-07-29T16:38:18","guid":{"rendered":"https:\/\/www.meslibellules.fr\/blog\/?p=25647"},"modified":"2025-07-29T18:38:20","modified_gmt":"2025-07-29T16:38:20","slug":"neurothemis-fluctuans","status":"publish","type":"post","link":"https:\/\/www.meslibellules.fr\/blog\/2025\/07\/29\/neurothemis-fluctuans\/","title":{"rendered":"Born\u00e9o, Sarawak &#8211; Neurothemis fluctuans (Fabricius, 1793)"},"content":{"rendered":"\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-2.webp\"><img decoding=\"async\" width=\"2211\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-2.webp\" alt=\"Neurothemis fluctuans m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025\" class=\"wp-image-25639 lazyload\" style=\"--smush-placeholder-width: 2211px; --smush-placeholder-aspect-ratio: 2211\/1504;width:476px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-2.webp 2211w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-2-768x522.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-2-1536x1045.webp 1536w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-2-2048x1393.webp 2048w\" data-sizes=\"(max-width: 2211px) 100vw, 2211px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Neurothemis fluctuans m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025<\/figcaption><\/figure>\n\n\n\n<p>Neurothemis fluctuans est un des trois Neurothemis au Sarawak. <br>Il se diff\u00e9rencie de N. ramburii et de N. terminata par sa petite taille, 22 \u00e0 25 mm (1), ses marques abdominales et sa nervation.<br>Ces d\u00e9tails sont mis en \u00e9vidence ci-dessous :<br>&#8211; le nombre de cellules dans le triangle de l&rsquo;aile ant\u00e9rieure est faible (cercle jaune), environ 4 \u00e0 7 (5 ci-dessous, 7 ci-dessus)<br>&#8211; de part et d&rsquo;autre de la car\u00e8ne dorsale abdominale, on trouve des paires de points (fl\u00e8ches blanches), plus ou moins discrets ; ces points ne sont pas visibles pour les deux autres esp\u00e8ces, la coloration \u00e9tant \u00e0 ce niveau fondue avec la tache de l&rsquo;apex des segments<br>&#8211; les marques lat\u00e9rales sur l&rsquo;abdomen (fl\u00e8ches jaunes) sont petites et centr\u00e9es dans les segments.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-1.webp\"><img decoding=\"async\" width=\"2155\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-1.webp\" alt=\"Neurothemis fluctuans m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025\" class=\"wp-image-25638 lazyload\" style=\"--smush-placeholder-width: 2155px; --smush-placeholder-aspect-ratio: 2155\/1504;width:495px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-1.webp 2155w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-1-768x536.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-1-1536x1072.webp 1536w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-1-2048x1429.webp 2048w\" data-sizes=\"(max-width: 2155px) 100vw, 2155px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Neurothemis fluctuans m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025<\/figcaption><\/figure>\n\n\n\n<p>La coloration alaire peut-\u00eatre proche de celle de <a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-ramburii-male-6.webp\">N. ramburii<\/a> (huit cellules dans le triangle alaire de ce sujet du Sarawak), tr\u00e8s variable \u00e9galement, avec la maculation de l&rsquo;aile post\u00e9rieure tr\u00e8s arrondie. Elle est par contre \u00e9loign\u00e9e de celle de <a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-terminata-male-1.webp\">N. terminata<\/a> (12 cellules dans le triangle de ce sujet du Sarawak) pour l&rsquo;aile post\u00e9rieure pour lequel elle ressemble \u00e0 celle de l&rsquo;aile ant\u00e9rieure, rejoignant le bord post\u00e9rieur de l&rsquo;aile en ligne droite.<\/p>\n\n\n\n<figure class=\"wp-block-gallery aligncenter has-nested-images columns-default is-cropped wp-block-gallery-1 is-layout-flex wp-block-gallery-is-layout-flex\">\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-3.webp\"><img decoding=\"async\" width=\"1593\" height=\"1504\" data-id=\"25640\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-3.webp\" alt=\"Neurothemis fluctuans m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025\" class=\"wp-image-25640 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-3.webp 1593w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-3-768x725.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-3-1536x1450.webp 1536w\" data-sizes=\"(max-width: 1593px) 100vw, 1593px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1593px; --smush-placeholder-aspect-ratio: 1593\/1504;\" \/><\/a><\/figure>\n\n\n\n<figure class=\"wp-block-image size-full\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-6.webp\"><img decoding=\"async\" width=\"1443\" height=\"1504\" data-id=\"25643\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-6.webp\" alt=\"Common Parasol male, Sarawak, Matang Wildlife Center, 03\/19\/2025\" class=\"wp-image-25643 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-6.webp 1443w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-6-768x800.webp 768w\" data-sizes=\"(max-width: 1443px) 100vw, 1443px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1443px; --smush-placeholder-aspect-ratio: 1443\/1504;\" \/><\/a><\/figure>\n<figcaption class=\"blocks-gallery-caption wp-element-caption\">Neurothemis fluctuans m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025 (6 cellules dans le triangle)<br> Matang Wildlife Center, 19\/03\/2025 (7 cellules dans le triangle)<\/figcaption><\/figure>\n\n\n\n<p>Seehausen et Dow (2) ont apport\u00e9 une r\u00e9vision tr\u00e8s attendue \u00e0 ce genre et on peut dans leur travail profiter de dessins et de reproductions explicites concernant tout d&rsquo;abord la coloration alaire ; on note comme N. fluctuans et ramburii peuvent \u00eatre ressemblants sur ce point, par exemple les variations N. fluctuans \u00ab\u00a0d\u00a0\u00bb et N. ramburii \u00ab\u00a0d\u00a0\u00bb et \u00ab\u00a0f\u00a0\u00bb, et qu&rsquo;on ne peut pas se fier \u00e0 ce seul \u00e9l\u00e9ment pour assoir l&rsquo;identification.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-large is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ailes-des-Neurothemis.webp\"><img decoding=\"async\" width=\"1622\" height=\"931\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ailes-des-Neurothemis.webp\" alt=\"Ailes des Neurothemis fluctuans, ramburii &amp; terminata, Seehausen &amp; Dow, 2016 (2)\" class=\"wp-image-25651 lazyload\" style=\"--smush-placeholder-width: 1622px; --smush-placeholder-aspect-ratio: 1622\/931;width:587px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ailes-des-Neurothemis.webp 1622w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ailes-des-Neurothemis-768x441.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/ailes-des-Neurothemis-1536x882.webp 1536w\" data-sizes=\"(max-width: 1622px) 100vw, 1622px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Ailes des Neurothemis fluctuans, ramburii &amp; terminata m\u00e2les<br> d&rsquo;apr\u00e8s Seenhausen et &amp; Dow, 2016 (2)<\/figcaption><\/figure>\n\n\n\n<p>Sur les abdomens, on note surtout sur la vue dorsale, les paires de points de N. fluctuans, alors que pour les deux autres esp\u00e8ces, ces points sont int\u00e9gr\u00e9s dans la tache sombre apicale.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><img decoding=\"async\" width=\"1300\" height=\"1204\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Abdomens-des-males-1.webp\" alt=\"Abdomens des m\u00e2les (1)\" class=\"wp-image-25656 lazyload\" style=\"--smush-placeholder-width: 1300px; --smush-placeholder-aspect-ratio: 1300\/1204;width:529px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Abdomens-des-males-1.webp 1300w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Abdomens-des-males-1-768x711.webp 768w\" data-sizes=\"(max-width: 1300px) 100vw, 1300px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><figcaption class=\"wp-element-caption\">Abdomens des m\u00e2les, d&rsquo;apr\u00e8s Seenhausen et Dow, 2016 (2)<\/figcaption><\/figure>\n\n\n\n<p>Dans le <em>Dragonflies of Japan<\/em> (3) on trouve le dessin ci-dessous qui met en \u00e9vidence les triangles alaires tr\u00e8s diff\u00e9rents entre N. fluctuans et ses deux cousins ramburii et terminata.<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Triangle-Neurothemis.webp\"><img decoding=\"async\" width=\"1000\" height=\"449\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Triangle-Neurothemis.webp\" alt=\"Triangle alaire des Neurothemis fluctuans, ramburii et terminata (3)\" class=\"wp-image-25611 lazyload\" style=\"--smush-placeholder-width: 1000px; --smush-placeholder-aspect-ratio: 1000\/449;width:694px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Triangle-Neurothemis.webp 1000w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Triangle-Neurothemis-768x345.webp 768w\" data-sizes=\"(max-width: 1000px) 100vw, 1000px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Triangle alaire des Neurothemis fluctuans, ramburii et terminata (3)<\/figcaption><\/figure>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-5.webp\"><img decoding=\"async\" width=\"1665\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-5.webp\" alt=\"Neurothemis fluctuans m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025\" class=\"wp-image-25642 lazyload\" style=\"--smush-placeholder-width: 1665px; --smush-placeholder-aspect-ratio: 1665\/1504;width:554px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-5.webp 1665w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-5-768x694.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-5-1536x1387.webp 1536w\" data-sizes=\"(max-width: 1665px) 100vw, 1665px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Neurothemis fluctuans m\u00e2le, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025 (4 cellules dans le triagle de l&rsquo;A.A.)<\/figcaption><\/figure>\n\n\n\n<p>C&rsquo;est une esp\u00e8ce qu&rsquo;on rencontre dans des habitats vari\u00e9s, pr\u00e8s ou \u00e0 distance de l&rsquo;eau. Il appr\u00e9cie les marais, les mares et les \u00e9tangs, mais aussi les rivi\u00e8res, les foss\u00e9s ou les canaux d&rsquo;irrigation pour peu que les milieux soient assez ouverts. On le rencontre le long des pistes foresti\u00e8res ou en bordure de for\u00eat.<br><br>Les femelles sont le plus souvent claires avec des marques sombres beaucoup plus visibles et le triangle alaire de leur aile ant\u00e9rieure ne comporte que deux cellules. On retrouve, de fa\u00e7on plus prononc\u00e9e encore, les marques abdominales caract\u00e9ristiques de l&rsquo;esp\u00e8ce (<a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Abdomens-des-femelles.webp\">comparaison des ailes<\/a> des trois esp\u00e8ces pr\u00e9sentent au Sarawak dans Seehausen et Dow, 2016(1)).<br>Si la femelle de gauche ci-dessous pourrait soulever quelques doutes quant aux marques lat\u00e9rales de son abdomen, on note la pr\u00e9sence des points d\u00e9j\u00e0 \u00e9voqu\u00e9s pour les m\u00e2les et le triangle \u00e0 deux cellules.<\/p>\n\n\n\n<figure class=\"wp-block-gallery aligncenter has-nested-images columns-default is-cropped wp-block-gallery-2 is-layout-flex wp-block-gallery-is-layout-flex\">\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-1.webp\"><img decoding=\"async\" width=\"1670\" height=\"1504\" data-id=\"25633\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-1.webp\" alt=\"Neurothemis fluctuans femelle, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025\" class=\"wp-image-25633 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-1.webp 1670w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-1-768x692.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-1-1536x1383.webp 1536w\" data-sizes=\"(max-width: 1670px) 100vw, 1670px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1670px; --smush-placeholder-aspect-ratio: 1670\/1504;\" \/><\/a><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-2.webp\"><img decoding=\"async\" width=\"1481\" height=\"1504\" data-id=\"25634\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-2.webp\" alt=\"Common Parasol female, Sarawak, Sama Jaya Nature Reserve, 03\/18\/2025\" class=\"wp-image-25634 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-2.webp 1481w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-2-768x780.webp 768w\" data-sizes=\"(max-width: 1481px) 100vw, 1481px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1481px; --smush-placeholder-aspect-ratio: 1481\/1504;\" \/><\/a><\/figure>\n<figcaption class=\"blocks-gallery-caption wp-element-caption\">Neurothemis fluctuans femelle, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025<\/figcaption><\/figure>\n\n\n\n<p>Il existe deux formes de femelles (2) ; l&rsquo;une commune aux ailes hyalines (parfois plus sombres \u00e0 l&rsquo;apex), l&rsquo;autre androchrome (isochromatic en anglais) dont la coloration du corps et des ailes est proche de celle des m\u00e2les et qui montre une nervation plus dense, avec 4 \u00e0 5 cellules dans le triangle de l&rsquo;aile ant\u00e9rieure.<\/p>\n\n\n\n<figure class=\"wp-block-gallery aligncenter has-nested-images columns-default is-cropped wp-block-gallery-3 is-layout-flex wp-block-gallery-is-layout-flex\">\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-4.webp\"><img decoding=\"async\" width=\"1609\" height=\"1204\" data-id=\"25636\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-4.webp\" alt=\"Neurothemis fluctuans femelle, Sarawak, Lorong Matang, 19\/03\/2025\" class=\"wp-image-25636 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-4.webp 1609w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-4-768x575.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-4-1536x1149.webp 1536w\" data-sizes=\"(max-width: 1609px) 100vw, 1609px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1609px; --smush-placeholder-aspect-ratio: 1609\/1204;\" \/><\/a><\/figure>\n\n\n\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-3.webp\"><img decoding=\"async\" width=\"2263\" height=\"1504\" data-id=\"25635\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-3.webp\" alt=\"Neurothemis fluctuans femelle, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025\" class=\"wp-image-25635 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-3.webp 2263w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-3-768x510.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-3-1536x1021.webp 1536w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-3-2048x1361.webp 2048w\" data-sizes=\"(max-width: 2263px) 100vw, 2263px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 2263px; --smush-placeholder-aspect-ratio: 2263\/1504;\" \/><\/a><\/figure>\n<figcaption class=\"blocks-gallery-caption wp-element-caption\">Neurothemis fluctuans femelle, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025<\/figcaption><\/figure>\n\n\n\n<p>Mais sur la photo ci-dessous, on serait bien en peine de compter le nombre de cellules du triangle de l&rsquo;aile ant\u00e9rieure de cette femelle \u00e0 l&rsquo;abdomen typique de l&rsquo;esp\u00e8ce : la nervation est compl\u00e8tement boulevers\u00e9e et d\u00e9form\u00e9e, \u00e0 l&rsquo;aile ant\u00e9rieure, comme \u00e0 l&rsquo;aile post\u00e9rieure. \u00c0 la fa\u00e7on dont les ailes sont pliss\u00e9es \u00e0 l&rsquo;apex, elle a sans doute connu des probl\u00e8mes d&rsquo;\u00e9mergence, mais elle ne semble pas s&rsquo;en soucier.<\/p>\n\n\n\n<figure class=\"wp-block-gallery aligncenter has-nested-images columns-default is-cropped wp-block-gallery-4 is-layout-flex wp-block-gallery-is-layout-flex\">\n<figure class=\"wp-block-image size-large\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4.webp\"><img decoding=\"async\" width=\"2189\" height=\"1504\" data-id=\"25641\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4.webp\" alt=\"Neurothemis fluctuans femelle, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025\" class=\"wp-image-25641 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4.webp 2189w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4-768x528.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4-1536x1055.webp 1536w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4-2048x1407.webp 2048w\" data-sizes=\"(max-width: 2189px) 100vw, 2189px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 2189px; --smush-placeholder-aspect-ratio: 2189\/1504;\" \/><\/a><\/figure>\n\n\n\n<figure class=\"wp-block-image size-full\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4bis.webp\"><img decoding=\"async\" width=\"1764\" height=\"1353\" data-id=\"25661\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4bis.webp\" alt=\"Neurothemis fluctuans femelle, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025\" class=\"wp-image-25661 lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4bis.webp 1764w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4bis-768x589.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-male-4bis-1536x1178.webp 1536w\" data-sizes=\"(max-width: 1764px) 100vw, 1764px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1764px; --smush-placeholder-aspect-ratio: 1764\/1353;\" \/><\/a><\/figure>\n<figcaption class=\"blocks-gallery-caption wp-element-caption\">Neurothemis fluctuans femelle, Sarawak, Sama Jaya Nature Reserve, 18\/03\/2025<\/figcaption><\/figure>\n\n\n\n<p>Sa distribution est large ; d&rsquo;ouest en est, on le donne en Inde, en raison de sa pr\u00e9sence dans les iles Nicobar et Adaman, jusqu&rsquo;\u00e0 Palawan, \u00e0 l&rsquo;ouest des Philippines, \u00e0 travers le sud du Myanmar, le nord de la Tha\u00eflande et Vietnam. Au sud, il est pr\u00e9sent sur la P\u00e9ninsule Malaise, Sumatra, Born\u00e9o et Java (2).<br><a href=\"https:\/\/www.iucnredlist.org\/species\/167488\/83383506\">IUCN Red List<\/a>.<br><br>Cette derni\u00e8re femelle a d\u00fb conna\u00eetre des moments difficiles ; c&rsquo;est sans doute une tentative de pr\u00e9dation qui lui a laiss\u00e9 un abdomen fortement d\u00e9sax\u00e9&#8230;<\/p>\n\n\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-5.webp\"><img decoding=\"async\" width=\"1915\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-5.webp\" alt=\"Common Parasol female, Sarawak, Mount Matang, 03\/23\/2025\" class=\"wp-image-25637 lazyload\" style=\"--smush-placeholder-width: 1915px; --smush-placeholder-aspect-ratio: 1915\/1504;width:519px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-5.webp 1915w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-5-768x603.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Neurothemis-fluctuans-femelle-5-1536x1206.webp 1536w\" data-sizes=\"(max-width: 1915px) 100vw, 1915px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Neurothemis fluctuans femelle, Sarawak, Mount Matang, 23\/03\/2025<\/figcaption><\/figure>\n\n\n\n<p><strong>\u00c9tymologie<\/strong><br>L&rsquo;esp\u00e8ce d\u00e9crite bri\u00e8vement par Fabricius en 1793 prend le nom de Polyneura palliata avec Rambur, 1842, puis Neurothemis fluctuans avec Brauer.<br><strong>Neurothemis<\/strong> (4), du grec neuro pour \u2013<em>nerf\u00a0<\/em>ou plut\u00f4t \u2013<em>nervure des ailes<\/em>, en entomologie, et de Themis \u2013 d\u00e9esse grecque de l\u2019ordre et de la justice, a \u00e9t\u00e9 utilis\u00e9 de nombreuses fois depuis Hagen (1861). Le nom de la d\u00e9esse de l\u2019ordre est particuli\u00e8rement bien adapt\u00e9 \u00e0 la taxonomie, science qui vise \u00e0 d\u00e9crire et ordonner les familles, genres et esp\u00e8ces. Brauer a cr\u00e9\u00e9 ce nom de genre en 1867 pour remplacer le genre Polyneura, cr\u00e9\u00e9 par Rambur en 1842, car ce dernier \u00e9tait d\u00e9j\u00e0 utilis\u00e9 chez les H\u00e9mipt\u00e8res. Ce nom de genre \u00e9tait destin\u00e9 \u00e0 insister sur le grand nombre de cellules de leurs ailes.<br><strong>Fluctuans<\/strong> ; Ngiam et Ng, 2022 (1) propose de traduire <em>fluctuans<\/em>, du verbe latin <em>fluctuare<\/em> qui signifie agit\u00e9 par les flots, ondulant, par le terme anglais de <em>fluctuating<\/em> qui signifie <em>fluctuant<\/em> pour la variabilit\u00e9 de la couleur du corps de l&rsquo;insecte qui va de brun rouill\u00e9 \u00e0 rouge brique. C&rsquo;est possible, mais d&rsquo;une part, je doute que Fabricius ait eu en main de nombreux  sujets au point de constater cette variabilit\u00e9, et d&rsquo;autre part, les sujets dont il a pu disposer ont mis des mois, si ce n&rsquo;est des ann\u00e9es \u00e0 lui parvenir et \u00e9taient donc dess\u00e9ch\u00e9s et d\u00e9color\u00e9s.<br>Une autre explication ; souvenons-nous que ce sont les balbutiements de l&rsquo;odonatologie, et il se pourrait que Fabricius se soit rendu compte ou ait voulu insister sur le fait que les ailes ne sont pas du tout planes, mais pr\u00e9sentent une succession de creux et de bosses, ce qui est beaucoup plus facilement visible sur des ailes sombres et color\u00e9es comme celles d&rsquo;un Neurothemis. Fluctuans serait alors proche de son sens original traduisant les ondulations de la surface de l&rsquo;aile (ce que j&rsquo;illustre ici avec une aile de <a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Aile-Calopteryx-virgo.webp\">Calopteryx virgo<\/a>).<br><br>Son nom vernaculaire anglais est Common parasol ; il est commun, c&rsquo;est vrai et il a pour habitude de se percher les ailes plus bas que l&rsquo;abdomen,  les ant\u00e9rieures tr\u00e8s basses, semblant abriter du soleil son thorax ou&#8230; ce qui est sous lui, comme un parasol.<\/p>\n\n\n\n<div style=\"height:30px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<p>1- Robin Ngiam &amp; Marcus Ng &#8211; A photographic guide to the Dragonflies and Damselflies of Singapore &#8211; John Beaufoy Publishing \u2013 2022<br>2- Malte Seehausen &amp; Rory A. Dow, 2016 \u2013\u00a0<a href=\"https:\/\/www.researchgate.net\/publication\/299553580_Morphological_studies_and_taxonomic_considerations_on_the_'reddishbrownwinged'_group_of_Neurothemis_Brauer_1867_with_the_description_of_N_taiwanensis_sp_nov_Odonata_Libellulidae#fullTextFileContent\">Morphological studies and taxonomic considerations on the \u2018reddishbrownwinged\u2019 group of Neurothemis Brauer, 1867 with the description of N. taiwanensis sp. nov. (Odonata: Libellulidae)<\/a>\u00a0\u2013 Journal of the International Dragonfly Fund.<br>3- Akira Ozono, Itsura Kawashima, Ryo Futahashi, 2012 \u2013 Dragonflies of Japan \u2013 Bun-ichi Sogo Shuppan Co.<br>4- <a href=\"https:\/\/www.researchgate.net\/publication\/283318421_The_Naming_of_Australia's_Dragonflies#fullTextFileContent\">The Naming of Australia\u2019s Dragonflies<\/a>, Ian Endersby &amp; Heinrich Fliedner, Busybird publishing, 2015.<\/p>\n\n\n\n<div style=\"height:31px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n\n\n<div class=\"wp-block-buttons is-vertical is-layout-flex wp-container-core-buttons-is-layout-fe9cc265 wp-block-buttons-is-layout-flex\">\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/blog\/odonates-du-sarawak\/\">Accueil Sarawak<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/divers\/liste-generale-odonates-du-site\/asia-odonates.php\">Listes des Odonates d&rsquo;Asie<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/pagesweb\/divers\/liste-generale-odonates-du-site\/all-odonates.php\">Listes continentales<\/a><\/div>\n<\/div>\n\n\n\n<p><\/p>\n","protected":false},"excerpt":{"rendered":"<p>Neurothemis fluctuans est un des trois Neurothemis au Sarawak. Il se diff\u00e9rencie de N. ramburii et de N. terminata par sa petite taille, 22 \u00e0 25 mm (1), ses marques abdominales et sa nervation.Ces d\u00e9tails sont mis en \u00e9vidence ci-dessous :&#8211; le nombre de cellules dans le triangle de l&rsquo;aile ant\u00e9rieure est faible (cercle jaune), [&hellip;]<\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"image","meta":{"site-sidebar-layout":"default","site-content-layout":"","ast-site-content-layout":"default","site-content-style":"default","site-sidebar-style":"default","ast-global-header-display":"","ast-banner-title-visibility":"","ast-main-header-display":"","ast-hfb-above-header-display":"","ast-hfb-below-header-display":"","ast-hfb-mobile-header-display":"","site-post-title":"","ast-breadcrumbs-content":"","ast-featured-img":"","footer-sml-layout":"","ast-disable-related-posts":"","theme-transparent-header-meta":"","adv-header-id-meta":"","stick-header-meta":"","header-above-stick-meta":"","header-main-stick-meta":"","header-below-stick-meta":"","astra-migrate-meta-layouts":"set","ast-page-background-enabled":"default","ast-page-background-meta":{"desktop":{"background-color":"var(--ast-global-color-4)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"ast-content-background-meta":{"desktop":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"footnotes":""},"categories":[1290],"tags":[],"class_list":["post-25647","post","type-post","status-publish","format-image","hentry","category-non-classe-fr","post_format-post-format-image"],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v27.5 - 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