{"id":28479,"date":"2026-02-08T18:51:15","date_gmt":"2026-02-08T17:51:15","guid":{"rendered":"https:\/\/www.meslibellules.fr\/blog\/2026\/02\/08\/ghana-tetrathemis-godiardi-lacroix-1921\/"},"modified":"2026-02-13T16:57:14","modified_gmt":"2026-02-13T15:57:14","slug":"ghana-tetrathemis-godiardi-lacroix-1921","status":"publish","type":"post","link":"https:\/\/www.meslibellules.fr\/blog\/2026\/02\/08\/ghana-tetrathemis-godiardi-lacroix-1921\/","title":{"rendered":"Ghana &#8211; Tetrathemis godiardi (Lacroix, 1921)"},"content":{"rendered":"\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-1.webp\"><img decoding=\"async\" width=\"1953\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-1.webp\" alt=\"Tetrathemis godiardi male, Ghana, Kakum N.P., 2026-01-14\" class=\"wp-image-28054 lazyload\" style=\"--smush-placeholder-width: 1953px; --smush-placeholder-aspect-ratio: 1953\/1504;aspect-ratio:1.298550724637681;width:502px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-1.webp 1953w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-1-768x591.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-1-1536x1183.webp 1536w\" data-sizes=\"(max-width: 1953px) 100vw, 1953px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Tetrathemis godiardi male, Ghana, Kakum N.P., 2026-01-14<\/figcaption><\/figure>\n\n<p>Tetrathemis godiardi, Libellulidae, is a rarity, very seldom encountered and photographed. Currently, only 4 or 5 photos can be found on the Web, on iNaturalist or observation.org.<br\/>It belongs to a genus that includes 14 species worldwide, found in Africa, Asia, and Oceania. Eight species are present in Africa, and these species might well belong to another genus, <em>Neophlebia<\/em> (which Selys created in 1869 for T. polleni).<br\/>Two species are present in Ghana, and we were fortunate enough to encounter them: this Tetrathemis godiardi and T. camerunensis, which is much more common.<br\/>These are charming small odonates due to their coloration and fabulous eyes, with very loose venation, quite similar from one species to another: for example, here are <a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-irregularis-male-1.webp\">T. irregularis<\/a> in Queensland (Australia) and <a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/2019\/06\/tetrathemis-platyptera-2.jpg\">T. platyptera<\/a> in Vietnam.<br\/>The specimen I am presenting first is young, immature, and its eyes are far from having their definitive color.  <\/p>\n\n<div class=\"wp-block-media-text is-stacked-on-mobile\"><figure class=\"wp-block-media-text__media\"><img decoding=\"async\" width=\"1398\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-3.webp\" alt=\"Tetrathemis godiardi male, Ghana, Kakum N.P., 2026-01-14\" class=\"wp-image-28056 size-full lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-3.webp 1398w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-3-768x826.webp 768w\" data-sizes=\"(max-width: 1398px) 100vw, 1398px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1398px; --smush-placeholder-aspect-ratio: 1398\/1504;\" \/><\/figure><div class=\"wp-block-media-text__content\">\n<p>African species are particularly similar (except for T. polleni) and differ almost exclusively by their anal appendages. There are only very slight differences, which could be considered individual variations, between this species and T. camerunensis; the spots on the abdomen are smaller, it does not have a clear mark on the sixth segment, as Lacroix (1921) tells us; but <a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-camerunensis-male-1.webp\">T. camerunensis<\/a> does not always have one either&#8230; <br\/><br\/>It is therefore necessary to compare the anal appendages, which I do below, and fortunately, they are very different.<\/p>\n\n\n\n<p><span style=\"font-size: 10pt;\">Tetrathemis godiardi male, Ghana, Kakum N.P., 2026-01-14<\/span><\/p>\n<\/div><\/div>\n\n<div style=\"height:32px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-appendices-anaux-1.webp\"><img decoding=\"async\" width=\"885\" height=\"465\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-appendices-anaux-1.webp\" alt=\"Tetrathemis godiardi male, anal appendages Ghana, Kakum N.P., 2026-01-14\" class=\"wp-image-28057 lazyload\" style=\"--smush-placeholder-width: 885px; --smush-placeholder-aspect-ratio: 885\/465;width:477px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-appendices-anaux-1.webp 885w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-appendices-anaux-1-768x404.webp 768w\" data-sizes=\"(max-width: 885px) 100vw, 885px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">T. godiardi male, anal appendages Ghana, Kakum N.P., 2026-01-14<\/figcaption><\/figure>\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-camerunensis-male-appendices-anaux.webp\"><img decoding=\"async\" width=\"883\" height=\"317\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-camerunensis-male-appendices-anaux.webp\" alt=\"Tetrathemis camerunensis male anal appendages, South Adansi, 2026-01-21\" class=\"wp-image-28061 lazyload\" style=\"--smush-placeholder-width: 883px; --smush-placeholder-aspect-ratio: 883\/317;width:473px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-camerunensis-male-appendices-anaux.webp 883w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-camerunensis-male-appendices-anaux-768x276.webp 768w\" data-sizes=\"(max-width: 883px) 100vw, 883px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">T. camerunensis male anal appendages, South Adansi, 2026-01-21<\/figcaption><\/figure>\n\n<p><\/p>\n\n<p>For Tetrathemis godiardi, it is noted that the anal appendages are short and the epiproct is longer than the cerci; for T. camerunensis, the cerci are long and divergent, concealing (in my photo) the shorter epiproct.<br\/>And I truly believe that photos are more illustrative than text, as Lacroix&rsquo;s description is quite obscure:<\/p>\n\n<figure class=\"wp-block-image aligncenter size-full\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Lacroix-1921-Tetrathemis-godiardi.webp\"><img decoding=\"async\" width=\"666\" height=\"109\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Lacroix-1921-Tetrathemis-godiardi.webp\" alt=\"Lacroix, 1921 - Tetrathemis godiardi\" class=\"wp-image-28069 lazyload\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 666px; --smush-placeholder-aspect-ratio: 666\/109;\" \/><\/a><figcaption class=\"wp-element-caption\">Lacroix, 1921 &#8211; Tetrathemis godiardi<\/figcaption><\/figure>\n\n<div style=\"height:32px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n<div class=\"wp-block-media-text has-media-on-the-right is-stacked-on-mobile\"><div class=\"wp-block-media-text__content\">\n<p><br\/>Lacroix specifies that the hindwing measures 24 mm and the abdomen 19 mm; the entire insect barely reaches 30 mm.<br\/><br\/>The species has been recorded from Liberia and Guinea to Cameroon.<br\/><a href=\"https:\/\/www.iucnredlist.org\/species\/158582\/5227297#geographic-range\">IUCN Red List<\/a>.<br\/><br\/><\/p>\n\n\n\n<p><span style=\"font-size: 10pt;\"><br>Tetrathemis godiardi male, Ghana, Kakum N.P., 2026-01-14<\/span><\/p>\n<\/div><figure class=\"wp-block-media-text__media\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-2.webp\"><img decoding=\"async\" width=\"1658\" height=\"1504\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-2.webp\" alt=\"Tetrathemis godiardi m\u00e2le, Ghana, Kakum N.P., 14\/01\/2026\" class=\"wp-image-28055 size-full lazyload\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-2.webp 1658w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-2-768x697.webp 768w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/Tetrathemis-godiardi-male-2-1536x1393.webp 1536w\" data-sizes=\"(max-width: 1658px) 100vw, 1658px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" style=\"--smush-placeholder-width: 1658px; --smush-placeholder-aspect-ratio: 1658\/1504;\" \/><\/a><\/figure><\/div>\n\n<div style=\"height:32px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n<p>Almost no information can be found on their biology or ecology; we encountered this immature male some distance from a pond. However, the mature specimen, shown below, was photographed (from afar!) on a small forest river where the current was slowed by rocks, forming a calmer basin; in the company of <em>Chlorocypha luminosa<\/em> and <em>radix,<\/em> <em>Elattoneura villiersi<\/em>, and <em>Allocnemis elongata<\/em>. <\/p>\n\n<figure class=\"wp-block-image aligncenter size-full is-resized\"><a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/t-godiardi.jpg\"><img decoding=\"async\" width=\"1328\" height=\"1025\" data-src=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/t-godiardi.jpg\" alt=\"Tetrathemis godiardi male, Ghana, Kakum N. P., 2026-01-13\" class=\"wp-image-28067 lazyload\" style=\"--smush-placeholder-width: 1328px; --smush-placeholder-aspect-ratio: 1328\/1025;aspect-ratio:1.2956330226246358;width:452px;height:auto\" data-srcset=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/t-godiardi.jpg 1328w, https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/t-godiardi-768x593.jpg 768w\" data-sizes=\"(max-width: 1328px) 100vw, 1328px\" src=\"data:image\/svg+xml;base64,PHN2ZyB3aWR0aD0iMSIgaGVpZ2h0PSIxIiB4bWxucz0iaHR0cDovL3d3dy53My5vcmcvMjAwMC9zdmciPjwvc3ZnPg==\" \/><\/a><figcaption class=\"wp-element-caption\">Tetrathemis godiardi male, Ghana, Kakum N. P., 2026-01-13 <\/figcaption><\/figure>\n\n<p><strong>Etymology<\/strong><br\/><strong>Tetrathemis<\/strong> (Brauer, 1868), from &#8211;<em>tetra<\/em> for &#8211;<em>four<\/em> and from <em>Themis<\/em> \u2013 Greek goddess of order and justice. The name of the goddess of order is particularly well suited to taxonomy, a science that aims to describe and classify families, genera, and species.<br\/>The number 4 refers, as Brauer describes, to the shape of the <em>cardinal<\/em> cell, which we perhaps improperly call a <em>triangle<\/em>, and which is quadrangular for this genus. <br\/><strong>Godiardi<\/strong> is a tribute to <a href=\"https:\/\/www.meslibellules.fr\/blog\/wp-content\/uploads\/image-3.png\">Maxime Godiard<\/a> who collected and transmitted this specimen from Ghana (Gold Coast). Little information is available about this colonial administrator, stationed at least in Senegal, who reportedly provided valuable logistical and human assistance to French naturalists on missions in French West Africa (AOF). <\/p>\n\n<div style=\"height:30px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n<p style=\"font-size:14px\"><strong>Ian Endersby &amp; Heinrich Fliedner<\/strong> &#8211; <a href=\"https:\/\/www.researchgate.net\/publication\/283318421_The_Naming_of_Australia's_Dragonflies#fullTextFileContent\">The Naming of Australia\u2019s Dragonflies<\/a>, Ian Endersby &amp; Heinrich Fliedner, Busybird publishing.<br\/><strong>Lacroix, 1921<\/strong> &#8211; <a href=\"https:\/\/www.biodiversitylibrary.org\/item\/44640#page\/836\/mode\/1up\">Deux Odonates nouvelles<\/a> &#8211; <em>Annales de la Soci\u00e9t\u00e9 entomologique de Belgique<\/em> (Vol. 61, Issue 10, p. 378).<\/p>\n\n<div style=\"height:30px\" aria-hidden=\"true\" class=\"wp-block-spacer\"><\/div>\n\n<div class=\"wp-block-buttons is-vertical is-layout-flex wp-container-core-buttons-is-layout-fe9cc265 wp-block-buttons-is-layout-flex\">\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/blog\/libellules-du-ghana-odonates-of-ghana\/\">Accueil Ghana<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"http:\/\/www.meslibellules.fr\/pagesweb\/divers\/liste-generale-odonates-du-site\/africa-odonates.php\">Liste africaine<\/a><\/div>\n\n\n\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link wp-element-button\" href=\"https:\/\/www.meslibellules.fr\/blog\/\">Accueil<\/a><\/div>\n<\/div>\n","protected":false},"excerpt":{"rendered":"<p>Tetrathemis godiardi, Libellulidae, is a rarity, very seldom encountered and photographed. Currently, only 4 or 5 photos can be found on the Web, on iNaturalist or observation.org.It belongs to a genus that includes 14 species worldwide, found in Africa, Asia, and Oceania. Eight species are present in Africa, and these species might well belong to [&hellip;]<\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"image","meta":{"site-sidebar-layout":"default","site-content-layout":"","ast-site-content-layout":"default","site-content-style":"default","site-sidebar-style":"default","ast-global-header-display":"","ast-banner-title-visibility":"","ast-main-header-display":"","ast-hfb-above-header-display":"","ast-hfb-below-header-display":"","ast-hfb-mobile-header-display":"","site-post-title":"","ast-breadcrumbs-content":"","ast-featured-img":"","footer-sml-layout":"","ast-disable-related-posts":"","theme-transparent-header-meta":"","adv-header-id-meta":"","stick-header-meta":"","header-above-stick-meta":"","header-main-stick-meta":"","header-below-stick-meta":"","astra-migrate-meta-layouts":"default","ast-page-background-enabled":"default","ast-page-background-meta":{"desktop":{"background-color":"var(--ast-global-color-4)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"ast-content-background-meta":{"desktop":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"tablet":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""},"mobile":{"background-color":"var(--ast-global-color-5)","background-image":"","background-repeat":"repeat","background-position":"center center","background-size":"auto","background-attachment":"scroll","background-type":"","background-media":"","overlay-type":"","overlay-color":"","overlay-opacity":"","overlay-gradient":""}},"footnotes":""},"categories":[38784,38792],"tags":[38793],"class_list":["post-28479","post","type-post","status-publish","format-image","hentry","category-ghana","category-libellulidae","tag-libellulidae-ghana","post_format-post-format-image"],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v27.3 - 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